Kazuya Kobayashi Department of Biology, Faculty of Agriculture and Life Science, Hirosaki University Hirosaki, Japan Takeshi KitanoDepartment of Biological Sciences, Graduate School of Science and Technology, Kumamoto UniversityKumamoto, Japan Yasuhiro Iwao Laboratory of Reproductive Developmental Biology, Division of Earth Sciences, Biology, and Chemistry, Graduate School of Sciences and Technology for Innovation, Yamaguchi UniversityYamaguchi, Japan Mariko Kondo Misaki Marine Biological Station, Graduate School of Science, The University of TokyoMiura, Japan